1. Department of Veterinary Internal Medicine, Graduate School of Agricultural and Life Sciences, The University of Tokyo,1 and
2. Japan Wildlife Research Center,2 Tokyo,
3. Fukuoka Municipal Zoo and Botanical Garden, Fukuoka,3
4. Department of Veterinary Internal Medicine, Faculty of Agriculture, University of Kagoshima, Kagoshima,4 and
5. Department of Pathobiology, Nihon University School of Veterinary Medicine, Kanagawa,5 Japan