1. The Scotch Whisky Research Institute, Research Avenue North, Riccarton, Edinburgh, EH14 4AP, Scotland
2. Department of Microbiology, University of Nigeria, Nsukka
3. Lake Chad Research Institute, Maiduguri, Borno State, Nigeria
4. International Centre for Brewing and Distilling, School of Life Sciences, Heriot-Watt University, Riccarton, Edinburgh, EH14 4AS, Scotland