1. The Scotch Whisky Research Institute, Research Avenue North, Riccarton, Edinburgh, EH14 4AP, Scotland
2. Department of Microbiology, University of Nigeria, Nsukka
3. Crop and Soil Systems Research Group, SRUC, West Mains Road, Edinburgh EH9 3JG, United Kingdom
4. International Centre for Brewing and Distilling, School of Life Sciences, Heriot-Watt University, Riccarton, Edinburgh, EH14 4AS, Scotland