1. Faculty of Agriculture, Ryukoku University, 1-5 Yokotani, Seta Oe-cho, Otsu 520-2194, Shiga, Japan
2. Research Center for Fermentation and Brewing, Ryukoku University, 1-5 Yokotani, Seta Oe-cho, Otsu 520-2194, Shiga, Japan
3. Nara National Research Institute for Cultural Properties, 2-9-1 Nijo, Nara 630-8577, Japan
4. Department of Archaeology, BioArCh, University of York, York YO10 5DD, UK