1. CAS Key Laboratory for Plant Biodiversity and Biogeography of East Asia Kunming Institute of Botany Chinese Academy of Sciences Kunming Yunnan650201China
2. Yunnan Key Laboratory for Integrative Conservation of Plant Species with Extremely Small Populations Kunming Institute of Botany Chinese Academy of Sciences Kunming Yunnan650201China
3. University of Chinese Academy of Sciences Beijing100049China
4. School of Resources and Environmental Engineering Anhui University Hefei Anhui province230601China
5. Department of Biosciences Durham University Stockton Road DurhamDH1 3LEUK