1. Kunming Botanical Garden, Kunming Institute of Botany; Chinese Academy of Sciences; Kunming 650204 China
2. Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany; Chinese Academy of Sciences; Kunming 650204 China
3. Chemi-ecology group of Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden; Chinese Academy of Sciences; Kunming 650201 China
4. School of Life Sciences; Yunnan University; Kunming 650200 China