1. State Key Laboratory of Phytochemistry and Plant Resources in West China, Kunming Institute of Botany Chinese Academy of Sciences Kunming China
2. University of Chinese Academy of Sciences Beijing China
3. CAS Key Laboratory for Plant Biodiversity and Biogeography of East Asia, Kunming Institute of Botany Chinese Academy of Sciences Kunming China
4. Yunnan Key Laboratory for Integrative Conservation of Plant Species with Extremely Small Populations Kunming China
5. Department of Animal and Plant Science University of Sheffield Sheffield UK