1. Kunming Botanical Garden, Kunming Institute of Botany; Chinese Academy of Sciences; Kunming 650201 China
2. Key Laboratory for Plant Diversity and Biogeography of East Asia; Kunming Institute of Botany; Chinese Academy of Sciences; Kunming 650201 China
3. College of Life Science and Technology; Honghe University; Mengzi 661199 China
4. Department of Botany and Biodiversity Research, Division of Systematic and Evolutionary Botany; University of Vienna; A-1030 Wien Rennweg 14 Austria
5. University of Chinese Academy of Sciences; Beijing 100049 China