1. Yunnan Key Laboratory for Integrative Conservation of Plant Species with Extremely Small Populations Kunming China
2. CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany Chinese Academy of Sciences Kunming China
3. College of Life Science Shenyang Normal University Shenyang China
4. School of Ecology and Environmental Science Yunnan University Kunming China
5. University of Chinese Academy of Sciences Beijing China
6. Ecology and Evolutionary Biology, School of Biosciences, Alfred Denny Building University of Sheffield Western Bank, Sheffield UK