1. Department of Integrative BiologyUniversity of Wisconsin‐MadisonMadison Wisconsin53706
2. Graduate School of Life ScienceHokkaido UniversitySapporo Hokkaido060‐0810 Japan
3. Department of PediatricsUniversity of Wisconsin‐MadisonMadison Wisconsin53706
4. Department of Ophthalmology and Visual SciencesUniversity of Wisconsin‐MadisonMadison Wisconsin53706