1. Key Laboratory of Plant Resources Conservation and Sustainable Utilization South China Botanical Garden Chinese Academy of Sciences Guangzhou 510650 China
2. University of the Chinese Academy of Sciences Beijing 100049 China
3. Department of Plant Biology Linnean Center for Plant Biology Uppsala BioCenter Swedish University of Agricultural Sciences Uppsala SE‐75007 Sweden
4. University of Bordeaux INRAE BIOGECO Cestas F‐33610 France
5. Center of Conservation Biology Core Botanical Gardens Chinese Academy of Sciences Guangzhou 510650 China