1. State Key Laboratory of Tree Genetics and Breeding, College of Biological Sciences and Technology Beijing Forestry University No. 35, Qinghua East Road Beijing 100083 China
2. National Engineering Research Center of Tree Breeding and Ecological Restoration, College of Biological Sciences and Technology Beijing Forestry University No. 35, Qinghua East Road Beijing 100083 China
3. Key Laboratory of Genetics and Breeding in Forest Trees and Ornamental Plants, Ministry of Education, College of Biological Sciences and Technology Beijing Forestry University No. 35, Qinghua East Road Beijing 100083 China
4. College of Forestry, State Forestry and Grassland Administration Key Laboratory of Silviculture in Downstream Areas of the Yellow River Shandong Agriculture University Taian Shandong 271018 China