1. Laboratory of Veterinary Microbiology, Faculty of Agriculture, Tottori University, 4-101 Koyama, Tottori 680-8553, Japan
2. The Avian Zoonosis Research Center, Faculty of Agriculture, Tottori University, 4-101 Koyama, Tottori 680-8553, Japan
3. Faculty of Veterinary Medicine, Khon Kaen University, 40002, Thailand