1. Graduate School of Engineering and Science, University of the Ryukyus, 1 Senbaru, Nishihara, Okinawa 903-0213, Japan
2. Department of Chemistry, Biology and Marine Science, Faculty of Science, University of the Ryukyus, 1 Senbaru, Nishihara, Okinawa 903-0213, Japan
3. Department of Natural History Sciences, Faculty of Science, Hokkaido University, North 10, West 8, Sapporo 060-0810 Japan
4. School of Animal, Plant & Environmental Science, University of the Witwatersrand, Private Bag 3, WITS 2050, South Africa