1. Graduate School of Engineering and Science, University of the Ryukyus, 1 Senbaru, Nishihara, Okinawa 903-0213, Japan
2. Sesoko Station, Tropical Biosphere Research Center, University of the Ryukyus Sesoko, Motobu, Okinawa 905-0227, Japan
3. Department of Chemistry, Biology and Marine Science, Faculty of Science, University of the Ryukyus, 1 Senbaru, Nishihara, Okinawa 903-0213, Japan