1. School of Animal, Plant & Environmental Sciences, University of the Witwatersrand, Private Bag 3 WITS, Johannesburg 2050, South Africa
2. Department of Chemistry, Biology and Marine Science, Faculty of Science, University of the Ryukyus, Nishihara, Okinawa, 903-0213, Japan
3. Department of Biological Sciences, Faculty of Science, Hokkaido University, North 10, West 8, Sapporo, 060-0810, Japan