1. State Key Laboratory of Plateau Ecology and Agriculture, Qinghai University, Xining, Qinghai, People’s Republic of China
2. Department of Agriculture and Forestry, College of Agriculture and Animal Husbandry, Qinghai University, Xining, Qinghai, People’s Republic of China
3. Qinghai Province Key Laboratory of Crop Molecular Breeding, Northwest Institute of Plateau Biology, Chinese Academy of Sciences, Xining, Qinghai, People’s Republic of China
4. Laboratory of Wheat Quality Improvement, The Innovative Academy of Seed Design, Chinese Academy of Sciences, Xining, Qinghai, People’s Republic of China