1. *Division of Pulmonary, Allergy and Critical Care, Perelman School of Medicine at the University of Pennsylvania, Philadelphia, PA;
2. †Department of Pathobiology, Wildlife Futures Program, University of Pennsylvania School of Veterinary Medicine, Kennett Square, PA;
3. ‡Bureau of Wildlife Management, Pennsylvania Game Commission, Harrisburg, PA; and
4. §Institute for Immunology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA