1. Kuju Agricultural Research Center, Faculty of Agriculture, Kyushu University 8780201, Japan
2. Muscle Biology and Meat Science laboratory, Research Faculty of Agriculture, Hokkaido University, 060-8589, Japan
3. Obihiro University of Agriculture and Veterinary Medicine, 080-8555, Japan
4. Laboratory of Animal Breeding and Genetics, Graduate School of Agricultural Science, Kobe University, 657-8501, Japan