1. CEFE, PSL-EPHE (Biogéographie et Ecologie des Vertébrés), CNRS, Univ. Montpellier, Univ Paul Valéry Montpellier, IRD, Montpellier, France
2. Department of Environmental Science, Faculty of Natural Resources, University of Tehran, Karaj, Iran
3. Department of Natural Resources Engineering (Habitats and Biodiversity), Faculty of Environment and Energy (FEE), Tehran Science and Research Branch, Islamic Azad University, Tehran, Iran
4. Department of Natural Resources, Isfahan University of Technology, Isfahan, Iran
5. Centre for Ecological Genomics and Wildlife Conservation, Department of Zoology, University of Johannesburg, Johannesburg, South Africa