1. School of Biological Sciences, Environment Centre Wales Bangor University Bangor LL57 2UW UK
2. Department of Biological Sciences The George Washington University 2023 G St. NW Washington DC 20052 USA
3. Cape Reptile Institute PO Box 1221 Oudtshoorn 6620 South Africa
4. Applied Biodiversity Research Division South African National Biodiversity Institute Claremont 7735 Cape Town South Africa
5. Department of Botany and Zoology University of Stellenbosch Private Bag X1 Matieland 7602 South Africa