1. Graduate School of Bioagricultural Sciences, Nagoya University, Furo-cho, Chikusa, Nagoya, Japan
2. Department of Horticulture, School of Horticulture and Plant Protection, Yangzhou University, Yangzhou, Jiangsu, P.R. China
3. NARO Tohoku Agricultural Research Center, Kariwano, Daisen, Akita, Japan
4. NARO Institute of Crop Science, Kannondai, Tsukuba, Ibaraki, Japan
5. School of Plant Biology, The University of Western Australia, Crawley, WA, Australia