1. Graduate School of Bioagricultural Sciences Nagoya University Furo‐Cho, Chikusa Nagoya 464‐8601 Japan
2. Department of Biotechnology, Graduate School of Engineering Osaka University 2‐1 Yamadaoka, Suita Osaka 565‐0871 Japan
3. Tohoku Agricultural Research Center National Agriculture and Food Research Organization Kariwano, Daisen Akita 019‐2112 Japan
4. Institute of Crop Science National Agriculture and Food Research Organization 2‐1‐2 Kannondai, Tsukuba Ibaraki 305‐8602 Japan
5. School of Plant Biology The University of Western Australia 35 Stirling Highway Crawley WA 6009 Australia