1. International Research Center for Agricultural & Environmental Biology, Faculty of Advanced Science and Technology, Kumamoto University, 2-39-1 Kurokami, Chuo Ward, Kumamoto 860-8555, Japan
2. L’Institut National de la Recherche Scientifique Centre Armand-Frappier Santé Biotechnologie (INRS-CAFSB), 531 des Prairies Blvd, Laval, QC H7V 1B7, Canada
3. Department of Botany, University of British Columbia, 6270 University Blvd, Vancouver, BC V6T 1Z4, Canada