1. Department of Genetics and Dominick Purpura Department of Neuroscience, Albert Einstein College of Medicine, Bronx, NY 1041, USA
2. Department of Biological Sciences, Howard Hughes Medical Institute, Columbia University, New York, NY 10027, USA
3. Department of Neuroscience and Developmental Biology, University of Vienna, Vienna 1090, Austria and
4. Research Institute of Molecular Pathology (IMP), Vienna Biocenter (VBC), Vienna 1030, Austria