1. Department of the Geophysical Sciences, University of Chicago, 5734 South Ellis Avenue, Chicago, IL 60637, USA
2. Senckenberg Biodiversity and Climate Research Center (BiK-F), Senckenberganlage 25, Frankfurt (Main) 60325, Germany
3. Section of Ecology, Behavior and Evolution, University of California San Diego, 9500 Gilman Drive, La Jolla, CA 92093-0116, USA