1. Negaunee Integrative Research Center, Field Museum, 1400 S. Lake Shore Drive, Chicago, IL 60605 USA
2. Natural History Museum, University of Oslo, Oslo, Norway
3. Department of Geography, Technical University of Liberec, Studentská 2, CZ-461 Liberec, Czech Republic
4. Centre for Ecological and Evolutionary Synthesis, Department of Biosciences, University of Oslo, Oslo, Norway