1. Kuroshio Science Program, Graduate School of Integrated Arts and Sciences, Kochi University, Nankoku, Kochi, Japan
2. Kuroshio Science Unit, Multidisciplinary Science Cluster, Kochi University, Nankoku, Kochi, Japan
3. Kuroshio Biological Research Foundation, Otsuki, Kochi, Japan
4. Marine Core Research Institute, Kochi University, Nankoku, Kochi, Japan