1. Department of Biology, University of Joensuu, PO Box 111, FIN-80101 Joensuu, Finland; Department of Geography and the Centre for Climate and Global Change Research, McGill University, Montreal, Quebec, H3A 2K6 Canada
2. Finnish Forest Research Institute, Unioninkatu 40 A, FIN-00170 Helsinki, Finland
3. Department of Biology, University of Joensuu, PO Box 111, FIN-80101 Joensuu, Finland
4. Finnish Forest Research Institute, PL 18, FIN-01301 Vantaa, Finland