1. CAS Key Laboratory of Coastal Environmental Processes and Ecological Remediation, Yantai Institute of Coastal Zone Research (YIC), Chinese Academy of Sciences (CAS); Shandong Key Laboratory of Coastal Environmental Processes, YICCAS, Yantai 264003, P. R. China
2. University of Chinese Academy of Sciences, Beijing 100049, P. R. China
3. Laboratory for Marine Fisheries Science and Food Production Processes, Qingdao National Laboratory for Marine Science and Technology, Qingdao 266237, P. R. China
4. Center for Ocean Mega-Science, Chinese Academy of Sciences, Qingdao 266071, P. R. China