1. 2Department of Biology, Gilman Lab, Dartmouth College, Hanover, New Hampshire 03755 USA;
2. 3Institute of Ecology, University of Georgia, Athens, Georgia 30602 USA;
3. 4Department of Plant, Soil and Insect Sciences and Graduate Program in Organismic and Evolutionary Biology, University of Massachusetts, Amherst, Massachusetts 01003 USA;
4. 5Department of Biology, Virginia Tech, Blacksburg, Virginia 24061 USA