1. Department of Plant Conservation, Federal Agency for Nature Conservation, Konstantinstr. 110, 53179 Bonn, Germany
2. Department of Biodiversity, Macroecology & Biogeography, University of Goettingen, Buesgenweg 1, 37077 Goettingen, Germany.
3. Department of Biology, University of Hawai‘i at Hilo, 200 W Kāwili Street, Hilo, HI 96720, USA.
4. USDA Forest Service, Pacific Southwest Research Station, Institute of Pacific Islands Forestry, 60 Nowelo Street,Hilo HI 96720, USA.