1. Department of Evolution, Ecology, and Organismal Biology, 1760 University Drive, The Ohio State University, Mansfield, Ohio 44906-1535 USA
2. Department of Biological Sciences, 331 Funchess Hall, Auburn University, Auburn, Alabama 36849-5407 USA
3. Department of Integrative Biology, 4184 Valley Life Sciences Building, University of California, Berkeley, Berkeley, California 94720-3140 USA