1. School of Biomedical Sciences, Faculty of Biological Sciences, University of Leeds, UK
2. Department of Biosciences, Section for Genetics and Evolutionary Biology, University of Oslo, Oslo, Norway
3. Molecular and Nanoscale Physics Group, School of Physics and Astronomy, University of Leeds, UK
4. Leeds Institute for Medical Research, St James' Teaching Hospital, Wellcome Trust Brenner Building, Leeds, LS9 7TF, UK