1. Department of Organismic Biology, Ecology and Evolution, University of California, Los Angeles, 900 Veteran Ave., Los Angeles, CA 90024‐1786, USA;
2. Departamento de Ciencias del Medio Natural, Universidad Publica de Navarra, ES–31006 Pamplona, Spain;
3. Division of Biology and Medicine, Brown University, Providence, RI 02912, USA