1. College of Urban and Environmental Sciences, Peking University, Beijing, 100871 China, State Key Lab of Environmental Aquatic Chemistry, Research Center for Eco-Environmental Sciences, Chinese Academy of Sciences, Beijing 100085, China, Research Institute of Genome-based Biofactory, National Institute of Advanced Industrial Science and Technology, 2-17-2-1, Tsukisamu-Higashi, Toyohira, Sapporo 062-8517, Hokkaido, Japan