1. Université de Nantes, CNRS, CEISAM UMR 6230, B.P. 92208, 2 rue de la Houssinière, 44322 Nantes Cedex 03, France
2. Department
of Fisheries and Wildlife, Michigan State University, East Lansing, Michigan 48824, United States
3. Institut Universitaire de France, Paris, 75005, France