1. School of Geography, Geology, and the Environment, Keele University, Staffordshire ST5 5BG, United Kingdom
2. School of Geographical Sciences, University of Bristol, Bristol BS8 1SS, United Kingdom
3. Chemical Ecology Group, School of Physical and Chemical Sciences, Keele University, Staffordshire ST5 5BG, United Kingdom
4. Department of Forest Ecology & Management, Swedish University of Agricultural Sciences, Umeå 750 07, Sweden
5. School of Geography, Earth, and Environmental Sciences, and Birmingham Institute of Forest Research, University of Birmingham, Birmingham B15 2TT, United Kingdom