1. College of Horticulture, Sanya Institute of Nanjing Agricultural University (NJAU), Nanjing 210095, China
2. Institute of Horticultural Crops, Xinjiang Academy of Agricultural Science, Urumqi,Xinjiang 830001, China
3. Institute of Botany, Jiangsu Province and Chinese Academy of Sciences, Nanjing 210014, China
4. The New Zealand Institute for Plant and Food Research Ltd (PFR), Auckland 92169, New Zealand