1. Key Laboratory of Horticultural Plant Biology, College of Horticulture and Forestry Science, Huazhong Agricultural University, Wuhan 430070, P.R. China
2. The New Zealand Institute for Plant and Food Research Ltd. (PFR), Auckland 1142, New Zealand
3. PFR, Palmerston North 4442, New Zealand
4. PFR, Te Puke 3182, New Zealand