1. MIVEGEC (UM1-UM2-CNRS 5290-IRD 224), Centre IRD, 911 avenue Agropolis BP 64501, 34394 Montpellier Cédex 5, France
2. Department of Ecology and Evolutionary Biology, Center for the Study of Complex Systems, University of Michigan, Ann Arbor, MI, USA
3. Fogarty International Center, National Institutes of Health, Bethesda, MD 20892, USA