1. Institut de Biologie de l'ENS (IBENS), Département de Biologie, École Normale Supérieure, CNRS, INSERM, Université PSL, 75005 Paris, France
2. Department of Ecology and Evolutionary Biology, Yale University, New Haven, CT 06511, USA
3. Department of Human Genetics, University of California, Los Angeles, 695 Charles E. Young Dr., Los Angeles, CA 90095, USA
4. Department of Biostatistics, University of California, Los Angeles, 695 Charles E. Young Dr., Los Angeles, CA 90095, USA
5. Center for Biodiversity and Global Change, Yale University, New Haven, CT 06511, USA