1. UMR LEMAR 6539 (UBO/CNRS/IRD/Ifremer), Ifremer, Technopôle Brest Iroise, BP 70, Plouzané 29280, France
2. Laboratoire de Génétique et Pathologie des Mollusques Marins, Ifremer—SG2M-LGPMM, Avenue Mus de Loup, La Tremblade 17390, France
3. Dyneco/BENTHOS, Ifremer, Technopôle Brest Iroise, BP 70, Plouzané 29280, France
4. Faculty of Veterinary Science, The University of Sydney, 425 Werombi Road, Camden, New South Wales 2570, Australia