1. Department of Ecology and Evolutionary Biology, Princeton University, Princeton, NJ, USA
2. Princeton School of Public and International Affairs, Princeton University, Princeton, NJ, USA
3. Mathematical Modelling of Infectious Diseases Unit, Institut Pasteur, CNRS, UMR2000, Paris, France
4. Santé publique France, French national public health agency, Saint Maurice, France
5. Department of Genetics, University of Cambridge, Cambridge, UK
6. Department of Biology, University of Florida, Gainesville, FL, USA
7. MRC–University of Glasgow Centre for Virus Research, Institute of Infection, Immunity and Inflammation, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow, UK