1. Academy of Agriculture and Forestry Sciences, Qinghai University, Xining 810016, China
2. Laboratory of Research and Utilization of Germplasm Resources in Qinghai-Tibet Plateau, Qinghai University, Xining 810016, China
3. Key Laboratory of Germplasm Resources Protection and Genetic Improvement of the Qinghai-Tibet Plateau in Ministry of Agriculture and Rural, Xining 810016, China
4. College of Agriculture and Animal Husbandry, Qinghai University, Xining 810016, China