1. State Key Laboratory of Desert and Oasis Ecology, Key Laboratory of Ecological Safety and Sustainable Development in Arid Lands, Xinjiang Institute of Ecology and Geography Chinese Academy of Sciences, Urumqi 830011, China
2. CAS Research Centre for Ecology and Environment of Central Asia, Urumqi 830011, China
3. Faculty of Geography and Environmental Sciences, Al-Farabi Kazakh National University, Almaty 050040, Kazakhstan
4. University of Chinese Academy of Sciences, Beijing 100049, China