1. Institute of Geography and Sustainability (IGD), Université de Lausanne, Mouline–Géopolis, 1015 Lausanne, Switzerland
2. Ecology, Department of Biodiversity and Environmental Management, Faculty of Biology and Environmental Sciences, Universidad de León, 24071 León, Spain
3. Centre d’Etudes Spatiales de la Biosphère, CESBIO, Université de Toulouse, CNES/CNRS/INRAE/IRD/UPS, 31000 Toulouse, France