1. Laboratory of Biotechnology Microbial Enzymatic and Biomolecules (LBMEB), Center of Biotechnology of Sfax, Sfax 3018, Tunisia
2. Laboratory of Enzyme Engineering and Microbiology, National School of Engineering of Sfax (ENIS), Sfax 1173-3038, Tunisia
3. Department of Life Sciences, Faculty of Science of Gabes, Gabès 6029, Tunisia
4. Laboratoire de Microbiologie Appliquée, Faculté des Sciences de la Nature et de la Vie, Université de Bejaia, Bejaia 06000, Algeria