1. Institute of Biotechnology, Addis Ababa University, Addis Ababa P.O. Box 1176, Ethiopia
2. Ethiopian Forest Development, P.O. Box 24536, Addis Ababa 1000, Ethiopia
3. Sustainable Forest Management Research Institute, University of Valladolid, Avda. Madrid 44, 34071 Palencia, Spain
4. Bio and Emerging Technology Institute, Addis Ababa P.O. Box 5954, Ethiopia