1. INRA, UMR 980 BAGAP, Joint Research Unit for Biodiversity, Agroecology, and Landscape Planning Rennes France
2. INRA, Agrocampus Ouest, UMR 985 ESE, Joint Research Unit for Ecosystem Ecology and Health Rennes France
3. Université Rennes 2, CNRS, UMR 6654 LETG, Joint Research Unit for Littoral Environments–Continental Environments–Remote Sensing–Geomatics Rennes France